Earl and Grady were able to kill so many Graboids in Tremors 2 because in the first Tremors, nobody had even see a Graboid before, let alone a small isolated town in the Midwest. They had to learn how to deal with a totally new enemy on the fly. In Tremors 2, Earl has already figured out a way to deal with the Graboids which is why he, Grady and Bert are able to kill so many of them.
Now it may seem like the Graboids were learning faster in Tremors, but in that case the creatures were all attacking one small community and often attacking the same targets. There were also less creatures, so Val and Earl had to use different tactics against the same Graboids. In Tremors 2, there's a lot more Graboids and Earl and Grady are killing them one at a time. So each time a Graboid encounters the RC-car trick, they're encountering it for the first time. They're also spread out over a wide area, so it's not like if one Graboid eats an RC-car and explodes his buddies right next to him figure out what happened. There's also how this time Bert, Grady, Earl and HQ were constantly in communication with each other so that they could easily find out how the others were doing.
Once the Shriekers start appearing, it becomes clear that Earl, Grady and Bert have no idea what to do initially. It's only because of Bert that they're able to figure out what makes the Shriekers tick, and even then they're still ill-equipped for the job. And come to think of it, since all the Graboids Earl and Grady encountered were getting ready to give birth to Shriekers, perhaps the Graboids kept going after the RC-cars because they needed food/sustenance much more than normal and were willing to forgo certain risks which caused many to be exploded by RC-cars.
This theme is even carried over into Tremors 3. Bert is able to kill an entire herd of Shriekers with a quad of 50 cal. machine guns because he was prepared. When the Ass-Blasters appear, the community again finds themselves unprepared and Bert inadvertently blows up his own house believing that it will stop them. However, once they figure out that the Ass-Blasters' innards are flammable and that they'll eat themselves into a food coma, the community once again easily gains the upper hand. Just look at El Blanco, Bert's dealt with Graboids for so long that one defective Graboid doesn't even phase him anymore.
Can't be too careful with all those weirdos running around.
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